Acharagma

From M&J Cactus Wiki
Acharagma
File:Acharagma roseanum.jpg
Kingdom Plantae
Family Cactaceae
Subfamily Cactoideae
Tribe Cacteae
SubTribe
Genus Acharagma
Species
Notes Taxonomy follows Joël Lodé (2015) with molecular support from Wallace (1995), Butterworth et al. (2002), Crozier (2005), Hernández-Hernández et al. (2011), and Vázquez-Sánchez et al. (2013).

Etymology

The name Acharagma means “without groove”, referring to the absence of an areolar groove on the tubercles, unlike related genera such as Coryphantha and Escobaria.

Description

Acharagma is a genus of small, globose to short cylindrical cacti with dense spination and reduced floral structures.

  • Habit: Solitary or forming small clumps (caespitose).
  • Roots: Fibrous to somewhat thickened.
  • Stems: Globose to short cylindrical, compact.
  • Ribs: Absent.
  • Tubercles: Prominent, without areolar groove and without nectar glands.
  • Areoles: Terminal, woolly when young.
  • Spines: Numerous, dense, covering the epidermis.
  • Flowers: Diurnal, apical, funnel-shaped; creamy white to yellow, often tinged pink or reddish; self-sterile.
  • Pollination: By insects.
  • Fruits: Naked, indehiscent; floral remnants persistent.
  • Seeds: Brown to black, finely foveolate.

Habitat

Occurs in restricted semi-desert environments:

  • xerophytic shrublands
  • rocky slopes and hillsides
  • limestone, sandstone, or gypsum substrates
  • cliff crevices and mountainous terrain

Often found among shrubs or in full sun, sometimes near seasonal watercourses.

Altitude range: approximately 1000–2650 m.

Distribution

Mexico:

  • Coahuila
  • Nuevo León

Taxonomy

The genus is well supported as distinct and monophyletic.

Key points:

  • Initially placed in Gymnocactus (Glass & Foster, 1970)
  • Later transferred to Turbinicarpus and Escobaria
  • Wallace (1995): molecular data indicated a distinct lineage
  • Glass & Zimmerman (1998): established genus status
  • Butterworth et al. (2002): separated from Escobaria
  • Crozier (2005) and Hernández-Hernández et al. (2011): confirmed distinct clade
  • Vázquez-Sánchez et al. (2013): confirmed monophyly

The genus is clearly separated from morphologically similar taxa.

Species

According to Joël Lodé (2015):

 ** Acharagma roseanum subsp. galeanense

Notes

  • A highly localized genus with very restricted distribution.
  • Morphologically similar to Escobaria, but lacking the diagnostic areolar groove.
  • The status of Acharagma roseanum subsp. galeanense remains uncertain and may warrant species rank.