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{| class="wikitable sortable" style="width:100%;"
== Classification of Cactaceae (McGinnie et al., 2025) ==
|+ '''Classification of the family Cactaceae (phylogenomic system by McGinnie et al., 2025)'''
|- style="background:#f2f2f2;"
! No. !! Subfamily !! Tribe !! Subtribe !! Genera
|-
| 1 || '''Leuenbergerioideae''' || || || ''Leuenbergeria''
|-
| 2 || '''Pereskioideae''' || || || ''Pereskia''
|-
| 3.1 || rowspan="3" | '''Opuntioideae''' || Opuntieae || || ''Airampoa, Brasiliopuntia, Consolea, Miqueliopuntia, Opuntia, Salmonopuntia, Tacinga''
|-
| 3.2 || Cylindropuntieae || || ''Cylindropuntia, Grusonia, Micropuntia, Pereskiopsis, Quiabentia''
|-
| 3.3 || Pterocacteae || || ''Austrocylindropuntia, Cumulopuntia, Maihueniopsis, Pterocactus, Tephrocactus''
|-
| 4 || '''Maihuenioideae''' || || || ''Maihuenia''
|-
| 5 || '''Blossfeldioideae''' || || || ''Blossfeldia''
|-
| 6.1 || rowspan="16" | '''Cactoideae''' || Lymanbensonieae || || ''Calymmanthium, Lymanbensonia''
|-
| 6.2 || Copiapoeae || || ''Copiapoa''
|-
| 6.3.1 || rowspan="3" | Cacteae || *Echinocactinae || ''Astrophytum, Aztekium, Echinocactus, Geohintonia, Sclerocactus''
|-
| 6.3.2 || Ferocactinae || ''Ferocactus, Glandulicactus, Kroenleinia, Leuchtenbergia, Stenocactus, Thelocactus''
|-
| 6.3.3 || Cactinae || ''Acharagma, Ariocarpus, Cochemiea, Coryphantha, Cumarinia, Epithelantha, Escobaria, Kadenicarpus, Lophophora, Mammillaria, Mammilloydia, Neolloydia, Obregonia, Oehmea, Ortegocactus, Pediocactus, Pelecyphora, Rapicactus, Strombocactus, Turbinicarpus''
|-
| 6.4.1 || rowspan="4" | Phyllocacteae || *Corryocactinae || ''Austrocactus, Brachycereus, Corryocactus, Eulychnia, Jasminocereus, Neoraimondia, Pfeiffera, Strophocactus''
|-
| 6.4.2 || Leptocereinae || ''Armatocereus, Castellanosia, Leptocereus''
|-
| 6.4.3 || Hylocereinae || ''Acanthocereus, Aporocactus, Disocactus, Epiphyllum, Hylocereus, Kimnachia, Pseudorhipsalis, Selenicereus, Weberocereus''
|-
| 6.4.4 || Echinocereinae || ''Backebergia, Bergerocactus, Carnegiea, Cephalocereus, Echinocereus, Escontria, Isolatocereus, Lemaireocereus, Lophocereus, Marginatocereus, Marshallocereus, Myrtillocactus, Nyctocereus, Pachycereus, Peniocereus, Polaskia, Pterocereus, Stenocereus''
|-
| 6.5 || Fraileeae || || ''Frailea''
|-
| 6.6 || Rhipsalideae || || ''Hatiora, Lepismium, Rhipsalidopsis, Rhipsalis, Schlumbergera''
|-
| 6.7 || Notocacteae || || ''Eriosyce, Neowerdermannia, Parodia, Yavia''
|-
| 6.8.1 || rowspan="7" | Cereeae || Uebelmanniinae || ''Uebelmannia''
|-
| 6.8.2 || Aylosterinae || ''Aylostera''
|-
| 6.8.3 || Rebutiinae || ''Browningia, Rebutia, Weingartia''
|-
| 6.8.4 || Gymnocalyciinae || ''Gymnocalycium''
|-
| 6.8.5 || Cereinae || ''Arrojadoa, Brasilicereus, Cereus, Cipocereus, Coleocephalocereus, Discocactus, Espostoopsis, Facheiroa, Floribunda, Lagenosocereus, Leocereus, Melocactus, Micranthocereus, Pierrebraunia, Pilosocereus, Praecereus, Siccobaccatus, Stephanocereus, Stetsonia, Xiquexique''
|-
| 6.8.6 || Reicheocactinae || ''Reicheocactus''
|-
| 6.8.7 || Trichocereinae || ''Acanthocalycium, Arthrocereus, Borzicactus, Cephalocleistocactus, Cleistocactus, Cremnocereus, Denmoza, Echinopsis, Espostoa, Haageocereus, Harrisia, Lasiocereus, Leucostele, Matucana, Mila, Oreocereus, Oroya, Pygmaeocereus, Rauhocereus, Samaipaticereus, Setiechinopsis, Vatricania, Weberbauerocereus, Winterocereus, Yungasocereus''
|}


<small>* Paraphyletic taxa are marked with an asterisk (*).</small>
The following hierarchical structure reflects the revised classification based on phylogenomic analysis of nuclear genes[cite: 4, 12, 106]. Paraphyletic taxa are indicated with an asterisk (*)[cite: 714].
 
* '''Subfamily Leuenbergerioideae''' [cite: 15, 701]
** ''Leuenbergeria'' [cite: 715, 857]
 
* '''Subfamily Pereskioideae''' [cite: 15, 701]
** ''Pereskia'' [cite: 715, 872]
 
* '''Subfamily Opuntioideae''' [cite: 15, 407, 701]
** Tribe Opuntieae [cite: 16, 407, 715]
*** ''Airampoa, Brasiliopuntia, Consolea, Miqueliopuntia, Opuntia, Salmonopuntia, Tacinga'' [cite: 715, 903-910]
** Tribe Cylindropuntieae [cite: 16, 407, 715]
*** ''Cylindropuntia, Grusonia, Micropuntia, Pereskiopsis, Quiabentia'' [cite: 715, 936-941]
** Tribe Pterocacteae [cite: 16, 407, 715]
*** ''Austrocylindropuntia, Cumulopuntia, Maihueniopsis, Pterocactus, Tephrocactus'' [cite: 715, 959-963]
 
* '''Subfamily Maihuenioideae''' [cite: 15, 701, 985]
** ''Maihuenia'' [cite: 715, 986]
 
* '''Subfamily Blossfeldioideae''' [cite: 15, 505, 701]
** ''Blossfeldia'' [cite: 715, 993]
 
* '''Subfamily Cactoideae''' [cite: 15, 701, 1000]
** Tribe Lymanbensonieae [cite: 16, 715, 808]
*** ''Calymmanthium, Lymanbensonia'' [cite: 715, 1019-1020]
** Tribe Copiapoeae [cite: 16, 715, 811]
*** ''Copiapoa'' [cite: 715, 1035]
** Tribe Cacteae [cite: 16, 715, 1044]
*** Subtribe *Echinocactinae [cite: 16, 715, 819]
**** ''Astrophytum, Aztekium, Echinocactus, Geohintonia, Sclerocactus'' [cite: 715, 1057-1059]
*** Subtribe Ferocactinae [cite: 16, 715, 1067]
**** ''Ferocactus, Glandulicactus, Kroenleinia, Leuchtenbergia, Stenocactus, Thelocactus'' [cite: 715, 1071-1074]
*** Subtribe Cactinae [cite: 16, 715, 1082]
**** ''Acharagma, Ariocarpus, Cochemiea, Coryphantha, Cumarinia, Epithelantha, Escobaria, Kadenicarpus, Lophophora, Mammillaria, Mammilloydia, Neolloydia, Obregonia, Oehmea, Ortegocactus, Pediocactus, Pelecyphora, Rapicactus, Strombocactus, Turbinicarpus'' [cite: 715, 1086-1107]
** Tribe Phyllocacteae [cite: 16, 715, 1163]
*** Subtribe *Corryocactinae [cite: 16, 715, 823]
**** ''Austrocactus, Brachycereus, Corryocactus, Eulychnia, Jasminocereus, Neoraimondia, Pfeiffera, Strophocactus'' [cite: 715, 1171-1180]
*** Subtribe Leptocereinae [cite: 16, 715, 825]
**** ''Armatocereus, Castellanosia, Leptocereus'' [cite: 715, 1212-1213]
*** Subtribe Hylocereinae [cite: 16, 715, 828]
**** ''Acanthocereus, Aporocactus, Disocactus, Epiphyllum, Hylocereus, Kimnachia, Pseudorhipsalis, Selenicereus, Weberocereus'' [cite: 715, 1231-1238]
*** Subtribe Echinocereinae [cite: 16, 715, 830]
**** ''Backebergia, Bergerocactus, Carnegiea, Cephalocereus, Echinocereus, Escontria, Isolatocereus, Lemaireocereus, Lophocereus, Marginatocereus, Marshallocereus, Myrtillocactus, Nyctocereus, Pachycereus, Peniocereus, Polaskia, Pterocereus, Stenocereus'' [cite: 715, 1285-1307, 1330-1335]
** Tribe Fraileeae [cite: 16, 715, 815]
*** ''Frailea'' [cite: 715, 1351]
** Tribe Rhipsalideae [cite: 16, 715, 1357]
*** ''Hatiora, Lepismium, Rhipsalidopsis, Rhipsalis, Schlumbergera'' [cite: 715, 1359-1362]
** Tribe Notocacteae [cite: 16, 715, 1378]
*** ''Eriosyce, Neowerdermannia, Parodia, Yavia'' [cite: 715, 1381-1387]
** Tribe Cereeae [cite: 16, 715, 1403]
*** Subtribe Uebelmanniinae [cite: 715, 1430]
**** ''Uebelmannia'' [cite: 715, 1433]
*** Subtribe Aylosterinae [cite: 16, 715, 1437]
**** ''Aylostera'' [cite: 715, 1439]
*** Subtribe Rebutiinae [cite: 16, 715, 1450]
**** ''Browningia, Rebutia, Weingartia'' [cite: 715, 1454-1456]
*** Subtribe Gymnocalyciinae [cite: 16, 715, 1464]
**** ''Gymnocalycium'' [cite: 715, 1466]
*** Subtribe Cereinae [cite: 16, 715, 1476]
**** ''Arrojadoa, Brasilicereus, Cereus, Cipocereus, Coleocephalocereus, Discocactus, Espostoopsis, Facheiroa, Floribunda, Lagenosocereus, Leocereus, Melocactus, Micranthocereus, Pierrebraunia, Pilosocereus, Praecereus, Siccobaccatus, Stephanocereus, Stetsonia, Xiquexique'' [cite: 722, 1529-1544]
*** Subtribe Reicheocactinae [cite: 16, 715, 1588]
**** ''Reicheocactus'' [cite: 715, 1591]
*** Subtribe Trichocereinae [cite: 16, 715, 1596]
**** ''Acanthocalycium, Arthrocereus, Borzicactus, Cephalocleistocactus, Cleistocactus, Cremnocereus, Denmoza, Echinopsis, Espostoa, Haageocereus, Harrisia, Lasiocereus, Leucostele, Matucana, Mila, Oreocereus, Oroya, Pygmaeocereus, Rauhocereus, Samaipaticereus, Setiechinopsis, Vatricania, Weberbauerocereus, Winterocereus, Yungasocereus'' [cite: 722, 1633-1634, 1644-1647, 1656-1662, 1676-1686]

Revision as of 10:02, 18 April 2026

Classification of Cactaceae (McGinnie et al., 2025)

The following hierarchical structure reflects the revised classification based on phylogenomic analysis of nuclear genes[cite: 4, 12, 106]. Paraphyletic taxa are indicated with an asterisk (*)[cite: 714].

  • Subfamily Leuenbergerioideae [cite: 15, 701]
    • Leuenbergeria [cite: 715, 857]
  • Subfamily Pereskioideae [cite: 15, 701]
    • Pereskia [cite: 715, 872]
  • Subfamily Opuntioideae [cite: 15, 407, 701]
    • Tribe Opuntieae [cite: 16, 407, 715]
      • Airampoa, Brasiliopuntia, Consolea, Miqueliopuntia, Opuntia, Salmonopuntia, Tacinga [cite: 715, 903-910]
    • Tribe Cylindropuntieae [cite: 16, 407, 715]
      • Cylindropuntia, Grusonia, Micropuntia, Pereskiopsis, Quiabentia [cite: 715, 936-941]
    • Tribe Pterocacteae [cite: 16, 407, 715]
      • Austrocylindropuntia, Cumulopuntia, Maihueniopsis, Pterocactus, Tephrocactus [cite: 715, 959-963]
  • Subfamily Maihuenioideae [cite: 15, 701, 985]
    • Maihuenia [cite: 715, 986]
  • Subfamily Blossfeldioideae [cite: 15, 505, 701]
    • Blossfeldia [cite: 715, 993]
  • Subfamily Cactoideae [cite: 15, 701, 1000]
    • Tribe Lymanbensonieae [cite: 16, 715, 808]
      • Calymmanthium, Lymanbensonia [cite: 715, 1019-1020]
    • Tribe Copiapoeae [cite: 16, 715, 811]
      • Copiapoa [cite: 715, 1035]
    • Tribe Cacteae [cite: 16, 715, 1044]
      • Subtribe *Echinocactinae [cite: 16, 715, 819]
        • Astrophytum, Aztekium, Echinocactus, Geohintonia, Sclerocactus [cite: 715, 1057-1059]
      • Subtribe Ferocactinae [cite: 16, 715, 1067]
        • Ferocactus, Glandulicactus, Kroenleinia, Leuchtenbergia, Stenocactus, Thelocactus [cite: 715, 1071-1074]
      • Subtribe Cactinae [cite: 16, 715, 1082]
        • Acharagma, Ariocarpus, Cochemiea, Coryphantha, Cumarinia, Epithelantha, Escobaria, Kadenicarpus, Lophophora, Mammillaria, Mammilloydia, Neolloydia, Obregonia, Oehmea, Ortegocactus, Pediocactus, Pelecyphora, Rapicactus, Strombocactus, Turbinicarpus [cite: 715, 1086-1107]
    • Tribe Phyllocacteae [cite: 16, 715, 1163]
      • Subtribe *Corryocactinae [cite: 16, 715, 823]
        • Austrocactus, Brachycereus, Corryocactus, Eulychnia, Jasminocereus, Neoraimondia, Pfeiffera, Strophocactus [cite: 715, 1171-1180]
      • Subtribe Leptocereinae [cite: 16, 715, 825]
        • Armatocereus, Castellanosia, Leptocereus [cite: 715, 1212-1213]
      • Subtribe Hylocereinae [cite: 16, 715, 828]
        • Acanthocereus, Aporocactus, Disocactus, Epiphyllum, Hylocereus, Kimnachia, Pseudorhipsalis, Selenicereus, Weberocereus [cite: 715, 1231-1238]
      • Subtribe Echinocereinae [cite: 16, 715, 830]
        • Backebergia, Bergerocactus, Carnegiea, Cephalocereus, Echinocereus, Escontria, Isolatocereus, Lemaireocereus, Lophocereus, Marginatocereus, Marshallocereus, Myrtillocactus, Nyctocereus, Pachycereus, Peniocereus, Polaskia, Pterocereus, Stenocereus [cite: 715, 1285-1307, 1330-1335]
    • Tribe Fraileeae [cite: 16, 715, 815]
      • Frailea [cite: 715, 1351]
    • Tribe Rhipsalideae [cite: 16, 715, 1357]
      • Hatiora, Lepismium, Rhipsalidopsis, Rhipsalis, Schlumbergera [cite: 715, 1359-1362]
    • Tribe Notocacteae [cite: 16, 715, 1378]
      • Eriosyce, Neowerdermannia, Parodia, Yavia [cite: 715, 1381-1387]
    • Tribe Cereeae [cite: 16, 715, 1403]
      • Subtribe Uebelmanniinae [cite: 715, 1430]
        • Uebelmannia [cite: 715, 1433]
      • Subtribe Aylosterinae [cite: 16, 715, 1437]
        • Aylostera [cite: 715, 1439]
      • Subtribe Rebutiinae [cite: 16, 715, 1450]
        • Browningia, Rebutia, Weingartia [cite: 715, 1454-1456]
      • Subtribe Gymnocalyciinae [cite: 16, 715, 1464]
        • Gymnocalycium [cite: 715, 1466]
      • Subtribe Cereinae [cite: 16, 715, 1476]
        • Arrojadoa, Brasilicereus, Cereus, Cipocereus, Coleocephalocereus, Discocactus, Espostoopsis, Facheiroa, Floribunda, Lagenosocereus, Leocereus, Melocactus, Micranthocereus, Pierrebraunia, Pilosocereus, Praecereus, Siccobaccatus, Stephanocereus, Stetsonia, Xiquexique [cite: 722, 1529-1544]
      • Subtribe Reicheocactinae [cite: 16, 715, 1588]
        • Reicheocactus [cite: 715, 1591]
      • Subtribe Trichocereinae [cite: 16, 715, 1596]
        • Acanthocalycium, Arthrocereus, Borzicactus, Cephalocleistocactus, Cleistocactus, Cremnocereus, Denmoza, Echinopsis, Espostoa, Haageocereus, Harrisia, Lasiocereus, Leucostele, Matucana, Mila, Oreocereus, Oroya, Pygmaeocereus, Rauhocereus, Samaipaticereus, Setiechinopsis, Vatricania, Weberbauerocereus, Winterocereus, Yungasocereus [cite: 722, 1633-1634, 1644-1647, 1656-1662, 1676-1686]
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